My article
Tønnessen, Morten 2001. Outline of an Uexkullian Bio-Ontology. Sign Systems Studies 29 (2): 683–691. With Russian ("Гранипы Юкскюлловской биоонтологии") and Estonian ("Uexkülli bio-ontoloogia piirjoni") abstract.
is referred to in
Marinakis, Yorgos 2012. On the semiosphere, revisited [p. 55 (Bibliography), Tønnessen 2001]. Signs vol. 6. Available online: https://tidsskrift.dk/index.php/signs/article/view/71478. 57 pp.
The reference occurs in the bibliography. I have not located any mention in the main body of text.
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