Tønnessen, M. (2011). Semiotics of being and Uexküllian phenomenology. In: Phenomenology/Ontopoiesis retrieving geo-cosmic horizons of antiquity (pp. 327–340). Springer Netherlands.
Tønnessen, M. (2016). Agency in biosemiotics and enactivism. In: Meaning, mind and communication: Explorations in cognitive semiotics. Peter Lang Publishing Group. pp. 69-82
are cited in
de Jesus, Paulo 2018. Thinking through enactive agency: sense-making, bio-semiosis and the ontologies of organismic worlds. Phenomenology and the Cognitive Sciences. Published online March 2nd 2018. DOI: https://doi.org/10.1007/s11097-018-9562-2.
Excerpts, p. 17:
BE [Biosemiotic Enactivism] endorses enactivism’s strong life-mind continuity but grounds it on Uexküll’s Umwelt theory and the bio-semiosis (sense-making/meaning-making) of living organisms (see also Meacham 2016).
In so doing BE explicitly distances itself from enactivism’s reliance on Hans Jonas and suggests in its place, following Morten Tønnessen (2011), an "Uexküllian phenomenology". Which, "is loyal not to Uexküll thought in detail but to the essential finding that nature is constituted by the intricate relations of all living creatures, which are all subjects of the phenomenal world at large" (Tønnessen 2011, p. 331). These relations are in turn conceived as bio-semiotic processes involving, at its most foundational-level, iconic (similarity-based) or/and indexical (causal) and at its most
developed-level symbolic (conventional) signs.
Biosemiotics, somewhat like enactivism, is still a field in its developing stages constituted by a loose group of connected yet different approaches. For an overview of the distinct schools and their relation see Barbieri (2009). For an exploration and comparison between enactive and biosemiotic approaches to agency see Tønnessen (2016).