Text on the evolutionary origin of Umwelt featured in Middlesex University Reading group session

My brief commentary article "The Evolutionary Origin(s) of the Umwelt. Biosemiotics" is included in this Spring´s readings for the reading group “Signs in Nature” at Middlesex University, along with Jablonka and Ginsburg´s Target article "Learning and the Evolution of Conscious Agents". This will happen in the session "Consciousness and Umwelt", which takes place Friday April 19th online.